| Publications: Parham, J.F., P.C.J. Donoghue, C.J. Bell, T.D. Calway, J.J. Head, P.A. Holroyd, J.G. Inoue, R.B. Irmis, W.G. Joyce, D.T. Ksepka, J.S.L. Patané, N.D. Smith, J.E. Tarver, M. Van Tuinen, Z. Yang, K.D. Angielczyk, J. Greenwood, C.A. Hipsley, L. Jacobs, P.J. Makovicky, J. Müller, K.T. Smith, J.M. Theodor, R.C.M. Warnock, and M.J. Benton. 2012. Best practices for justifying fossil calibrations. Systematic Biology 61(2):346-359. 
Wheatley, P.V., Hofman, A.E., Holroyd, P.A., Goodwin, M.B., and Brown, S.T. 2011. Calcium isotope fractionations in vertebrates from modern and fossil ecosystems. Geological Society of America, Annual Meeting, Minneapolis, MN
Archibald, J.D., W. A. Clemens, K. Padian, T. Rowe, N. Macleod, P. M. Barrett, A. Gale, P. Holroyd, H.-D. Sues, N. C. Arens, J. R. Horner, G. P. Wilson, M. B. Goodwin, C. A. Brochu, D. L. Lofgren, S. H. Hurlbert, J. H. Hartman, D. A. Eberth, P. B. Wignall, P. J. Currie, A. Weil, G. V. R. Prasad, L. Dingus, V. Courtillot, A. Milner, A. Milner, S. Bajpai, D. J. Ward, and A. Sahni. 2010. Cretaceous Extinctions: Multiple Causes. Science. 328: 973
Gunnell, G.F., P.D. Gingerich and P.A. Holroyd. 2010. Ptolemaiida. Pp.
83-88 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa,
University of California Press.
Holroyd, P.A. 2010. Macroscelidea. Pp. 89-98 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.
Holroyd, P.A. 2010. Tubulidentata. Pp. 107-112 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California
Press.
Holroyd, P.A., F. Lihoreau, G. F. Gunnell, and E. Miller. 2010. Anthracotheriidae. Pp. 843-852 in L. Werdelin and W.J. Sanders (eds.), Cenozoic Mammals of Africa, University of California Press.
Dirks, W., R.L. Anemone, P.A. Holroyd, D.J. Reid, and P. Walton. 2009. Phylogeny, life history, and the timing of molar crown formation in two archaic ungulates, Meniscotherium and Phenacodus (Mammalia, 'Condylarthra'). Pp. 3-8 in T. Koppe, G. Meyer, and K.W. Alt (eds.), Comparative Dental Morphology. Frontiers of Oral Biology 13. Karger Press, Basel.
Goodwin, M.B., T. Deméré, P. Holroyd, R. Wilson, and S. Dowker. 2009. Unusual preservation of fossil baleen (Cetacea: Mysticeti) from the Miocene of California, USA: Comparative morphology and stable isotope evidence for seasonality and growth. Journal of Vertebrate Paleontology 29(3):107A.
Grossman, A. and P.A. Holroyd. 2009. Miosengi butleri, gen. et sp. nov., (Macroscelidea) from the Kalodirr Member, Lothidok Formation, early Miocene of Kenya. Journal of Vertebrate Paleontology 29(3):957-960.
Holroyd, P. A. 2009. Past records of Elephantulus and Macroscelides: geographic and taxonomic issues. Afrotherian Conservation 7:3-7.
Holroyd, P. A. and N.J. Stevens. 2009. Differentiation of Phiomys andrewsi from Lavocatomys aequatorialis (n. gen., n. sp.) (Rodentia: Thryonomyoidea) in the Oligo-Miocene interval on continental Africa. Journal of Vertebrate Paleontology. 29(4):1331-1334.
Stevens, N.J., P.A. Holroyd, E.M. Roberts, P.M. O'Connor, and M.D. Gottfried. 2009. Kahawamys mbeyaensis (n. gen., n. sp.) (Rodentia: Thryonomyoidea) from the Late Oligocene Rukwa Rift Basin, Tanzania. Journal of Vertebrate Paleontology 29(2):631-634.
Bourque, J., J. Hutchison, P.A. Holroyd, and J. Bloch. 2008. A new
kinosternoid (Testudines: Dermatemydidae) from the Paleocene-Eocene
boundary of the Bighorn Basin, Wyoming and its paleoclimatological
implications. Journal of Vertebrate Paleontology 27(3 Suppl.):55A.
Clementz, M. A., P. A. Holroyd, and P. L. Koch. 2008. Identifying aquatic habits of herbivorous mammals through stable isotope analysis. Palaios 23(9):574-585.
Clementz, M. T. and P. A. Holroyd. 2008. From river horses to sea cows: using stable isotopes in tooth enamel to identify semiaquatic mammals in the fossil record. The 6th International Conference on Applications of Stable Isotope Techniques to Ecological Studies. Honolulu, HI August 25-29.
Clementz, M. T., and P. A. Holroyd. 2008. Isotopic modeling of the hippo ecomorph: identifying aquatic habits of herbivorous mammals through stable isotope analysis. The 6th International Conference on Applications of Stable Isotope Techniques to Ecological Studies. Honolulu, HI August 25-29.
Clementz, M. T., P. A. Holroyd, and P. L. Koch. 2008. Identifying aquatic
habits of herbivorous mammals through stable isotope analysis. Palaios
23(9): 547-585
Dirks, W., R. L. Anemone, P. A. Holroyd, D. J. Reid, and P. Walton. 2008. Phylogeny, life history and the timing of crown formation in two archaic
ungulates, Meniscotherium and Phenacodus (Mammalia, “Condylarthra”), 14th International Symposium on Dental Morphology, Greifswald, Germany,
August 27-30, Abstracts with Programs, p.O8
Goodwin, M. B. and P. A. Holroyd. 2008. Fractionation in fossil bioapatite: phylogeny as a confounding factor. Journal of Vertebrate Paleontology 28(3):85A
Head, J.J., and P.A. Holroyd. 2008. Assembly and biogeography of North
American Paleogene snake faunas based on an expanded fossil record.
Journal of Vertebrate Paleontology 27(3 Suppl.):90A.
Holroyd, P.A. & Strait, S.G. 2008. New data on Loveina (Primates: Omomyidae) from the early Eocene Wasatch Formation and implications for washakiin relationships, pp. 243-257, in J.G. Fleagle and C.C. Gilbert, Elwyn Simons: A Search for Origins. Springer.
Holroyd, P.A. 2008. New data on dental eruption patterns in condylarths
and afrotheres. Journal of Vertebrate Paleontology 27(3 Suppl.):93A
Smith, K. T., B.-A. S. Bhullar, and P. A. Holroyd. 2008. Earliest African
record of the Varanus stem-clade (Squamata: Varanidae) from the earliest
Oligocene of Egypt. Journal of Vertebrate Paleontology 28(3): 909-913.
Clementz, M., and P. Holroyd. 2007. Using stable isotope analysis ot test inferred semi-aquatic habits in fossil herbivores from the early Oligocene of Egypt. Journal of Vertebrate Paleontology 27(3):59A.
Goodwin, M.B., P.G. Grant, G. Bench, and P. Holroyd. 2007. Elemental composition and diagenetic alteration of dinosaur bone: Distinguishing micron-scale spatial and compositional heterogeneity using PIXE. Palaeogeography, Palaeoclimatology, Palaeoecology 253:458-476. doi:10.1016/j.palaeo.2007.06.017.
Holroyd, P., and G. Gunnell. 2007. New data on Anthracotheriidae (Artiodactyla) from the Paleogene of Egypt. Journal of Vertebrate Paleontology 27(3):90A.
Holroyd, P.A., T. Tsubamoto, N. Egi, R. L. Ciochon, M. Takai, Soe Thura Tun, Chit Sein, and G.F. Gunnell. 2006. A rhinocerotid perissodactyl from the late middle Eocene Pondaung Formation, Myanmar. Journal of Vertebrate Paleontology 26(2):491-494.
Egi, N., P.A. Holroyd, T. Tsubamoto, Aung Naing Soe, M. Takai, and R.L. Ciochon. 2005. Proviverrine hyaenodontids (Creodonta: Mammalia) from the Eocene of Myanmar and phylogenetic analysis of the proviverrines from the Para-Tethys area. Journal of Systematic Paleontology 3(4):337-358.
Head, J.J., Holroyd, P.A., Hutchison J.H., and R.L. Ciochon. 2005. First report of snakes (Serpentes) from the late middle Eocene Pondaung Formation, Myanmar. Journal of Vertebrate Paleontology 25(1):246-250.
Holroyd, P.A., and J.C. Mussell. 2005. Origin and relationships of the Macroscelidea and Tubulidentata. Pp. 71-83 in K.D. Rose and J.D. Archibald (eds.), The Rise of Placental Mammals: Origins and Relationships of the Major Extant Clades. Johns Hopkins University Press, Baltimore, Maryland.
Holroyd, P.A., J.F. Parham, and J.H. Hutchison. 2005. A reappraisal of some Paleogene turtles from the southeastern United States. Journal of Vertebrate Paleontology 25(4):979-982.
Stidham, T.A., P.A. Holroyd, G.F. Gunnell, R.L. Ciochon, T. Tsubamoto, N. Egi, and M. Takai. 2005. An ibis-like bird (Aves: cf. Threskiornithidae) from the late middle Eocene of Myanmar. Contributions from the University of Michigan Museum of Paleontology 31(7):179-184.
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