I have devoted my whole life to the study of Nature, and yet a single sentence may express all that I have done. I have shown that there is a correspondence between the succession of Fishes in geological times and the different stages of their growth in the egg, -- that is all. It chanced to be a result that was found to apply to other groups and has led to other conclusions of a like nature.Louis Agassiz, 1869
Biography of Agassiz
The son of a minister, Jean Louis Rodolphe Agassiz was born on May 28, 1807 in the village of Montier, in the French-speaking part of Switzerland. Agassiz was educated in the universities of Switzerland and Germany as a physician, like many naturalists of the time. He studied with prominent German biologists, including Oken and Döllinger. These men were followers of Naturphilosophie, a German Romantic philosophy that sought for metaphysical correspondences and interconnections within the world of living things. Though Agassiz later renounced this philosophy, he was never quite able to free himself from its influence. Receiving his medical degree from the University of Erlangen in 1830, he went to Paris in November 1831 to study comparative anatomy under Cuvier, the most famous naturalist in Europe. Cuvier was so impressed with Agassiz's work on fossil fish that he turned over to Agassiz his own notes and drawings for a planned work on fossil fish. Cuvier died in April 1832, yet although their relationship lasted only six months, Agassiz always considered himself an intellectual heir of Cuvier For the rest of his life, Agassiz promoted and defended Cuvier's geological catastrophism and classification of the animals. With the publication of his vast work on the fossil record of fish, Poissons fossiles, Agassiz's reputation began to grow in the scientific community.
After Cuvier's death, Agassiz took up a professorship at the Lyceum of Neuchatel in Switzerland, where for thirteen years he worked on many projects in paleontology, systematics, and glaciology. Agassiz took up the study of glaciers in 1836 as something of a sideline, but his contributions made him known as the "Father of Glaciology." Observing the glaciers of his native Switzerland, Agassiz noticed the marks that glaciers left on the Earth: great valleys; large glacial erratic boulders carried long distances; scratches and smoothing of rocks; mounds of debris called moraines pushed up by glacial advances. He realized that in many places these signs of glaciation could be seen where no glaciers existed. Previous scientists had variously explained these features as made by icebergs or floods; Agassiz integrated all these facts to formulate his theory that a great Ice Age had once gripped the Earth, and published his theory in Étude sur les glaciers in 1840. His later book, Système glaciare (1847), presented further evidence for his theory, gathered all over Europe: Agassiz later found even more evidence of glaciation in North America.
In 1846, Agassiz came to the United States; in 1848 he accepted a professorship at Harvard. He immediately set about organizing and acquiring funding for a great museum of natural history. In 1859 his dream came true with the founding of the Museum of Comparative Zoology, which opened its doors in 1860. Agassiz labored for support of science in his adopted homeland; he and his colleagues urged the creation of a National Academy of Sciences, and Agassiz became a founding member in 1863. Agassiz was also appointed a regent of the Smithsonian Institution in 1863. He campaigned constantly for funds and resources for American science, and for his research projects in particular -- and the funding grew and grew (although it never seemed quite enough for all that Agassiz wanted to do -- and although Agassiz himself never quite finished most of his grand projects).
Agassiz's Scientific Thought
Agassiz stayed loyal to Cuvier's classification, which divided the animal kingdom into four branches: Vertebrata, Insecta, Vermes (worms) and Radiata (radially symmetrical animals). Within each embranchement the classes could be ranked from lowest to highest; the orders in each class could be similarly ranked, and so on down to the species level, with Homo sapiens sitting at the very top of the scale of life. The cornerstone of Agassiz's biological thought was his belief that the gradation from low to high forms, in any taxon, paralleled the order of appearance in the fossil record, the order of stages in the organisms' development, and the distribution and ecology of the taxon. The "lowest" forms were found lowest in the rock record, earliest in embryonic development, and at the highest latitudes. Agassiz summed up his thought in his Essay on Classification, first published in 1851:
. . . the phenomena of animal life correspond to one another, whether we compare their rank as determined by structural complication with the phases of their growth, or with their succession in past geological ages; whether we compare this succession with their relative growth, or all these different relations with each other and with the geographical distribution of animals upon the earth. The same series everywhere!Darwin, and many others after him, accepted these parallelisms as providing evidence for evolution. Darwin wrote in The Origin of Species that "this doctrine of Agassiz accords well with the theory of natural selection," and Haeckel in particular invoked the "recapitulation of phylogeny by ontogeny" in support of evolution. But Agassiz was no evolutionist; in fact, he was probably the last reputable scientist to reject evolution outright for any length of time after the publication of The Origin of Species. Agassiz saw the Divine Plan of God everywhere in nature, and could not reconcile himself to a theory that did not invoke design. He defined a species as "a thought of God." As he wrote in his Essay on Classification:
The combination in time and space of all these thoughtful conceptions exhibits not only thought, it shows also premeditation, power, wisdom, greatness, prescience, omniscience, providence. In one word, all these facts in their natural connection proclaim aloud the One God, whom man may know, adore, and love; and Natural History must in good time become the analysis of the thoughts of the Creator of the Universe. . .Agassiz continued Cuvier's catastrophism theory -- the Earth had been periodically wracked by global catastrophes, after each of which new species of animals and plants had appeared. Followers of Cuvier had suggested that the Biblical Flood was the last catastrophe. Agassiz replaced the Flood with his glaciers, which he thought had been formed instantaneously all over the world; he called glaciers "God's great plough," and tried unsucessfully to find evidence of glaciation in Brazil.
Agassiz's works on living and fossil fish and on glaciers have remained classics. His work on glaciers revolutionized geology, and drove another nail in the coffin of the Biblical Flood as a serious scientific hypothesis. He trained and influenced a generation of American zoologists and paleontologists, including Alpheus Hyatt, William Healey Dall, David Starr Jordan, Nathaniel Shaler, and Edward S. Morse. He left a mark on the development and the practice of American science, and brought science to "the man in the street" as no one else had before. People from all over the world read his books, sent him specimens, and asked his advice. By the time of his death, on December 14, 1873, he was publicly recognized as America's leading scientist.
His philosophy of nature, aiming to understand the Divine Plan, is the last great expression of the old school of natural theology, started by men like John Ray almost two hundred years before. Natural theology had once inspired countless scientists, including Darwin and his forerunners, but by the time of publication of the Origin of Species it had largely run out of steam, unable to offer any real explanation for natural phenomena except "God made it that way." Within Agassiz's lifetime, and much to his grief, most of his students -- including his son Alexander, a well-known naturalist in his own right -- became evolutionists, though not necessarily Darwinians.
Yet Agassiz still made lasting contributions to evolutionary biology and systematics. His construction of a classification that did not depend on any process of evolution has been followed up in the work of the "pattern cladist" school of systematists,who try to reconstruct organismal relationships without relying on assumptions of what processes generated them. His finding of parallels between ontogeny, paleontology, and morphology was rapidly adopted by biologists like Haeckel and used to support evolution. Today, these parallels are known not to be exact correspondences, but the links between development and evolution remain an area of active research. Perhaps Agassiz's greatest lasting insight was the realization that paleontology, embryology, ecology, and biogeography had to contribute to any classification that purported to show the true relationships of organisms -- even if those relationships, to Agassiz, existed only in the mind of God. As he wrote in his Essay on Classification:
Classification seems to me to rest upon too narrow a foundation when it is chiefly based on structure. Animals are linked together as closely by their mode of development, by their relative standing in their respective classes, by the order in which they have made their appearance upon earth, by their geographical distribution, and generally by their connection with the world in which they live, as by their anatomy. All these relations should, therefore, be fully expressed in a natural classification; and though structure furnishes the most direct indication of some of these relations, always appreciable under every circumstance, other considerations should not be neglected which may complete our insight into the general plan of creation.
The 1869 French translation of Agassiz's Essay on classification includes a chapter, not found in earlier editions, that clearly explains Agassiz's reasons for resisting Darwinism. This chapter is available on the Web thanks to its translator, Paul J. Morris.